The Brexit Party was the clear winner in the UK European election results, with the pro-EU Lib Dems coming second.
The Conservatives and Labour suffered heavy losses, with the former heading for less than 10% of the vote.
Brexit Party leader Nigel Farage said the two main parties “could learn a big message” from the results.
Overall, out of 64 MEPs declared so far, Mr Farage’s party has won 28, the Lib Dems 15, Labour 10, Greens seven, the Tories three and Plaid Cymru one.
All 28 EU member states are electing MEPs. According to early results, the main centre-right and centre-left blocs in the European Parliament have lost ground, amid a surge in support for Eurosceptic parties.
The UK had been due to leave the EU on 29 March, but when that deadline was missed, participation in the election became mandatory.
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The Brexit Party topped the polls in every region of England apart from London. It also dominated in Wales, with Plaid Cymru second.
Scotland is yet to formally declare, but with most votes counted, the SNP is on course for a resounding victory, with about 38% of the vote.
Meanwhile, Northern Ireland’s count does not begin until Monday, with the results expected on Tuesday.
Polling expert Sir John Curtice said the results showed just how polarised the country had become.
BBC political editor Laura Kuenssberg said it was the worst performance for the Conservatives as a party going back to 1832.
“If this was a first-past-the-post election, they would not have taken a single seat,” she said.
- The UK is electing 73 MEPs and 10 out of 12 regions have so far declared
- The Brexit Party has received the highest share of the vote in nine of the 10 regions – it is heading for first place overall, with about a third of the vote
- The Conservative Party has been widely rejected by the electorate and is heading for fifth place
- The Lib Dems have taken second place with 20% of the vote – a vote that is clearly much higher in areas that backed Remain. In London, the Lib Dems came first and the Tories fifth
- The Green Party has also made gains, seeing its best performance since 1989
- Newly-formed Change UK has not won any seats, while UKIP has also haemorrhaged support to The Brexit Party – both will likely gain about 3% of the vote
- The Labour Party has fallen to third place overall – fifth in Scotland – and is on course to end up with less than 15% of the vote, an even worse performance than the previous low in 2009
Mr Farage said if the UK did not leave the EU by the new deadline of 31 October, these results would be repeated at a general election.
“Never before in British politics has a party just six weeks old won a national election,” he tweeted.
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Brexiteer and Conservative MEP for the South East Daniel Hannan – one of only three Tory MEPs elected so far – told the BBC it was his party’s “worst ever result”.
“We voted to leave (the EU) and we haven’t left – it’s that simple,” he said.
Tory leadership hopeful and former Foreign Secretary Boris Johnson said the public had delivered “a crushing rebuke” to both major parties for failing to deliver Brexit.
Labour leader Jeremy Corbyn placed blame for that failure squarely with the Conservatives, arguing it meant the elections had “become a proxy second referendum”.
He said his party would “reflect on these results on both sides of the Brexit divide” and the issue “will have to go back to the people” through a general election or a public vote.
Labour’s deputy leader Tom Watson said the party needed to “urgently” re-think its Brexit position after the “disastrous” election results.
Alastair Campbell, former spin doctor to Tony Blair and prominent member of the People’s Vote campaign for another referendum, said for the first time in his life he had voted for the Lib Dems.
In contrast to the two biggest parties, smaller ones with an unambiguous message on Brexit were rewarded.
Lib Dem Sir Ed Davey said his party was “leading the Remain vote” and had “an awful lot to be pleased about”.
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Green Party co-leader Sian Berry said the established parties had been rejected “so badly” and the public were “crying out for something new – they’ve seen that in the Green Party”.
She insisted the results were “not a victory” for The Brexit Party as voters had also shown support for Remain parties.
Change UK leader Heidi Allen told the BBC her party – newly formed from ex-Labour and Tory MPs – was “down, but we are not out”.
It’s probably best read as approximating to a draw – between the Remain and Leave sides, that is. (I’m leaving out the nationalists.)
Two parties, both in favour of leaving without a deal overwhelmingly almost entirely supported by Leave voters: The Brexit Party and UKIP. Total vote: 35%
Three parties: the Liberal Democrats, the Greens and Change UK, all in favour of another referendum. Total vote: 35%.
I think the oddest truth is that what this election has done is two things.
One is to demonstrate just how polarised the public are on this issue.
The second is to remind us, as the opinion polls have been telling us, that support for Remain and support for Leave are still very close to each other.
The highest Brexit Party votes were recorded in Castle Point, in Essex, (59%), and Boston (56%) and South Holland (55%), both in Lincolnshire.
The best Lib Dem performances occurred in Gibraltar (77%), Richmond upon Thames (52%) and Kingston upon Thames (47%).
The best Green performances were in Brighton and Hove (36%), Bristol (35%) and Stroud (28%).
Turnout was just below 37%. That puts it on course to be the second highest in any European election – second only to 2004 when the turnout was partly inflated by the use of all-postal ballots in some parts of England.
Places that voted most strongly to Remain in the referendum saw turnout increase more compared with the last election in 2014 than places which voted most heavily for Leave.
Both of our big main parties are being punished severely for the meltdown in Westminster over Brexit.
The flip-side for the smaller parties? The Lib Dems are buoyant and of course, Nigel Farage is back with a vengeance.
A European election may not be a direct proxy for a UK general election, but nevertheless, after tonight, millions of voters in regions up and down the country will be represented by parties other than the big two.
It also casts doubt on the idea that compromise can win out. Instead, the clear message seems to be that people want clarity – whether that be Leave or Remain – when it comes to Brexit.
Leavers want to Leave, Remainers want to Remain, and efforts to persuade people they should back something in the middle have failed.